Education Background:

• March 1955 Graduated from Hiroshima University, majoring in botany
• March 1960 Completed required courses for a DSc. in plant ecology, Graduate School, Hiroshima University
• February 1962 Awarded D.Sc.

Academic career:

• April 1967 Associate professor in the Faculty of Liberal Arts, Nagasaki University
• July 1970 Professor, Faculty of Liberal Arts, Nagasaki University
• October 1997 Professor, Faculty of Environmental Studies, Nagasaki University
• March 1998 Reached retirement age at Nagasaki University
• May 1998 Awarded the title of Professor Emeritus at Nagasaki University
• Publications (mainly related to Galapagos and island biology)

Book:

• The Galapagos Islands - the Cradle of Evolution Theory. 194 pages. Chuko New Book 91, CHUOKORON, Inc. (in Japanese)
• Island Life (translation from Island Life by J. Sparks). 142 pages, Kodansha, Ltd. (in Japanese)
• The Galapagos Islands - the Cradle of Evolution Theory, new edition. 212 pages. Chuko New Book 690. CHUOKORON, Inc. (in Japanese)
• The Galapagos Islands - Evolution and Ecology of Organisms. 80 pages. Iwanami Graphics 28, Iwanami Shoten, Publishers. (in Japanese)
• Natural History of Island Plants. 246 pages. Kodansha Meche, Kodansha Ltd. (in Japanese)
• Galapagos Islands - World Heritage, Ecotourism, and El Nino. 257 pages, Kadokawa Selected Book 340, Kadokawa Shoten Publishing Co., Ltd. (in Japanese)
• Vegetation of Oceanic Islands - Galapagos Archipelago. In (Shuzo Ito, ed.) Pacific Islands. Weekly Asahi, Plant World Encyclopedia - No. 107/. 262-266 pages. Asahi Shimbun Company. (in Japanese)

Scientific paper:

• 1966. Ecological approaches to the vegetation of the Galapagos Islands. Noticias de Galapagos 12:10-12.
• 1971. A study of vegetation in the Galapagos Islands. Noticias de Galapagos 17:10-13.
• 1974. Fens and bogs in the Galapagos Islands. Hikobia 7:39-52. (Co-author, D. Weber)
• 1975. Effects of rain-shadow on vegetation distribution in the Galapagos Islands. XIII Pacific Science Congress, Vancouver, Canada.
• 1988. Species diversity of mainland- and island forests in the Pacific area. Vegetatio 77:193-200.
• 1988. Population structure, stand-level dieback and recovery of Scalesia pedunculata forest in the Galapagos Islands. Ecological Research 3:333-339. (Co-author, D. Mueller-Dombois)
• 1990. Herbaceous and ericaceous communities in the highland of Santa Cruz, the Galapagos Islands. Monog. Syst. Missouri Bot. Gard. 32:47-58.
• 1992. Altitudinal change in plant endemism and species turn-over and diversity on Isla Santa Cruz, the Galapagos Islands. Pacific Science. 46(2):251-268.
• 1995 Phytogeography and ecology of Scalesia (Compositae) endemic to the Galapagos Islands. Pacific Science 49: 17-30.
• 1997. List of plant specimens collected in the Galapagos Islands, Ecuador. Bull. Fac. Liberal Arts, Nagasak Univ. (Nat. Sci.), 38(1): 53-144.
• 1999. Aboveground biomass and soil nutrient pools of a Scalesia pedunculata montane forest on Santa Cruz, Galapagos. Ecological Research, 14:405-408. (Co-author, K. Kitayama)
• 2003. Zonation pattern, succession process and invasion by aliens in species-poor insular vegetation of the Galapagos Islands. Global Environmental Research, 7 (1) : 39-58.